Louis Rees-Zammit will make his first start for Wales in more than two years when they take on Japan in a crucial Autumn international on Saturday.
The Penarth-born wing marked his return to international rugby when he came off the bench during last Sunday's 52-28 defeat against Argentina in Cardiff.
Head coach Steve Tandy has named four changes as Wales aim for their first win on home soil since 2023 - and with seedings at stake ahead of next month's draw for the 2027 Rugby World Cup.
As expected, Dewi Lake takes over the captaincy from Ospreys teammate Jac Morgan, who will miss the rest of the Autumn Nations Series after dislocating his shoulder.
Leicester Tigers flanker Olly Cracknell, who made his Test debut last weekend, is promoted to number eight, with Aaron Wainwright at blind-side flanker and Alex Mann shifting to number seven.
Props Nicky Smith and Archie Griffin come in for Rhys Carré and Keiron Assiratti, who drop to the bench with Tom Rogers missing out altogether.
Tandy said: "The props started the game really well and played 45 minutes. We were always planning to rotate those round after the six-day turnaround. Then there’s a reshuffle in the back row: Alex Mann goes to seven."
"We feel he’s got all the qualities to play six or seven. Some of his attacking ability you’ve seen on the weekend and his defensive energy was outstanding. Then bringing Olly Cracknell in at No. 8 and moving Aaron Wainwright to six gives us a really good balance in the back row."
"Louis Rees-Zammit comes in on the wing. He played a good few minutes last weekend and we feel he’s ready to return to the starting line-up."
“I think we’ve definitely seen lots of how we want to play against Argentina. There was lots to be pleased about – our attack I thought was excellent and I saw a lot of physicality – but there’s lots to improve. So that’s what we’ve been focusing on. We’re looking to build on the first performance into this weekend."
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